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Inbreeding in Chefs-de-Race, Their Sires and Their Dams
Jones and Bogart, in "Genetics of the Horse" (Edwards Brothers, Inc., Ann Arbor, Michigan, 1971), define inbreeding as the mating of two individuals more closely related than the average of the breed. They highly recommend that serious breeders know the amount of inbreeding in their stock in order to appreciate the rate at which homozygosity is developing, homozygosity being the condition where genes inherited from the sire and the dam are alike. The amount of inbreeding estimates the percentage of genes put in a homozygous condition. This estimate is always less than the actual because records will not allow for every relationship that exists. Horses that are highly homozygous tend to stamp their own characteristics on their offspring, a phenomenon often called "prepotency". Prepotency also may be considered a regular or predictable transmission of particular traits.
The amount of inbreeding can be calculated using a formula proposed by Wright in 1923 (Mendelian Analysis of the Pure Breeds of Livestock, J. Hered. 14:339-348). Wright's formula generates a fraction (the coefficient of inbreeding) which, when multiplied by 100, affords the amount of inbreeding as a percentage. Wright's equation is:

where FX is the inbreeding coefficient of the horse in question, FA is the inbreeding coefficient of the common ancestor, n1 is the number of generations from the sire to the common ancestor, and n2 is the number of generations from the dam to the common ancestor.
The information below includes Wright's Six-Generation Inbreeding Coefficient (6-Gen IC) expressed as a percentage for all 217 chefs-de-race as well as for their sires and their dams. The raw data are preceded by some general statistics.
Observations:
The general trend of inbreeding percentage among chefs-de-race declined steadily from the late 19th century until it began to level off among chefs-de-race born after the mid-20th century. This trend is displayed in the chart on the left. The approximate overall rate at which inbreeding among chefs-de-race has declined since the late 19th century until the present time is 0.019 percent per year, although as noted, the decline is non-linear. Whether the pattern found among chefs-de-race represents the pattern found among all Thoroughbreds is not known. The chart below on the right shows the decrease in inbreeding among chefs-de-race born within discreet time frames (before 1920; 1920 to 1929; 1930 to 1939; 1940 to 1949; 1950 to 1959; 1960 to 1969 and since 1969) using the median 6-Gen IC within each time segment. Here we use the median figure because the sample sets are small and the median deemphasizes data points very far from the average. This avoids a situation in which one outlying data point can significantly skew the average. The chart shows a similar pattern for annual leading earners since 1902, although the slope is flatter and corresponds to an annual decrease in inbreeding among these runners of 0.007 percent per year. Also, within every time frame, inbreeding among chefs-de-race is greater thah inbreeding among the runners.
The amount of inbreeding found among chefs-de-race covers the range from over 14% for Ultimus down to 0% for Speak John, with some of the most influential stallions being found at both ends of the spectrum. For example, Havresac II (a key link to Ribot) and Bayardo (a key link to Hyperion) are among the most highly inbred, while Native Dancer (a close up ancestor of Raise a Native, Mr. Prospector and Northern Dancer) is among the least inbred.
Arguably, the best runners among the chefs-de-race are Secretariat (0.24%), Man o' War (1.13%), Ribot (0.69%), Nearco (1.95%) and Sea-Bird (0.49%). All except Nearco are inbred below the average or median percentage of inbreeding for the time frame in which they were foaled.
There is no significant pattern covering the relationship between inbreeding in the chefs-de-race and inbreeding in their parents. Most (82, 37.8%) are more inbred than one parent while 67 (30.9%) are less inbred than either parent and 64 (29.5%) are more inbred than either parent. The remaining 4 (1.8%) are inbred the same as one parent. The following charts, again using median 6-Gen ICs for each time period, displays a similar decline in the percentage of inbreeding over time for the chefs-de-race, their sires and their dams, with the decline rates being 0.018% per year for chefs-de-race, 0.020% per year for their sires and 0.023% per year for their dams.
For 58.5% of the chefs-de-race (127), their sire is more highly inbred than their dam. For 40.6% of the chefs-de-race (88), the reverse is true. There are two examples (0.9%) where the sire and dam of the chef-de-race exhibit the same level of inbreeding.
There are observable differences in inbreeding among chefs-de-race sorted by aptitudinal contribution; however, these differences are not statistically significant.
The data clearly show that inbreeding among chefs-de-race and high end performers has decreased over the last century. What is not clear at this time is whether the greater inbreeding among chefs-de-race compared to the runners actually contributes to their ability to transmit consistent aptitudinal type to their descendants. It also is not clear whether the decrease in inbreeding among chefs-de-race has leveled off permanently in recent years or is a temporary situation. Note, however, that for both chefs-de-race and for the runners, the median 6-Gen IC has increased for those foaled between 1960 and 1969 and those foaled afterwards.
General Statistics:
Average 6-Gen IC of all chefs-de-race:
1.64%
Average 6-Gen IC of the sires of all chefs-de-race: 1.76%
Average 6-Gen IC of the dams of all chefs-de-race: 1.65%
% Inbreeding over Time
Average 6-Gen IC of chefs-de-race born 1893-1919: 2.62%
Average 6-Gen IC of chefs-de-race born 1920-1929: 2.16%
Average 6-Gen IC of chefs-de-race born 1930-1939: 1.70%
Average 6-Gen IC of chefs-de-race born 1940-1949: 1.50%
Average 6-Gen IC of chefs-de-race born 1950-1959: 1.24%
Average 6-Gen IC of chefs-de-race born 1960-1969: 1.18%
Average 6-Gen IC of chefs-de-race born 1970-1997: 1.30%
|
Chefs-de-Race with
the Highest 6-Gen IC Ultimus: 14.32% Havresac II: 12.71% Bayardo: 5.13% Heliopolis: 4.91% Turn-to: 4.64% Colorado: 4.52% Mossborough: 4.51% In Reality: 4.50% Broad Brush: 4.45% The Tetrarch: 4.35% |
Chefs-de-Race with
the Lowest 6-Gen IC Speak John: 0.00% Native Dancer: 0.05% Reliance II: 0.05% Bold Bidder: 0.10% King's Bishop: 0.10% Relko: 0.10% Stage Door Johnny:0.10% Roman: 0.20% My Babu: 0.20% Lost Code: 0.20% Prince John: 0.20% Promised Land: 0.20% |
Inbreeding in Chefs-de-Race Relative to Their Sire and Dam
Chefs-de-race with 6-Gen IC < 6-Gen IC
of either sire or dam: 67 (30.9%)
Chefs-de-race with 6-Gen IC > 6-Gen IC of either sire or dam: 64 (29.5%)
Chefs-de-race with 6-Gen IC > 6-Gen IC in sire and < 6-Gen IC in dam: 34
(15.7%)
Chefs-de-race with 6-Gen IC < 6-Gen IC in sire and > 6-Gen IC in dam: 48
(22.1%)
Chefs-de-race with 6-Gen IC = 6-Gen IC in either sire or dam: 4 (1.8%)
% Inbreeding in Chefs-de-Race by Aptitudinal Group Representation
Average 6-Gen IC for chefs-de-race in Brilliant: 1.74%
Average 6-Gen IC for chefs-de-race in Intermediate: 1.64%
Average 6-Gen IC for chefs-de-race in Classic: 1.40%
Average 6-Gen IC for chefs-de-race in Solid: 1.41%
Average 6-Gen IC for chefs-de-race in Professional: 1.60%
| CHEF-DE-RACED | YEAR | SIRE | 6-GEN IC | DAM | 6-GEN IC | |||
| A.P. INDY | 1989 | 3.33% | SEATTLE SLEW | 1.57% | WEEKEND SURPRISE | 3.57% | ||
| ABERNANT | 1946 | 0.49% | OWEN TUDOR | 3.72% | RUSTOM MAHAL | 0.15% | ||
| ACK ACK | 1966 | 0.89% | BATTLE JOINED | 1.76% | FAST TURN | 1.71% | ||
| ADMIRAL DRAKE | 1931 | 1.38% | CRAIG AN ERAN | 1.22% | PLUCKY LIEGE | 1.37% | ||
| ALCANTARA II | 1908 | 1.47% | PERTH | 2.99% | TOISON D'OR | 2.02% | ||
| ALIBHAI | 1938 | 1.63% | HYPERION | 3.72% | TERESINA | 0.68% | ||
| ALIZIER | 1947 | 0.34% | TELEFERIQUE | 1.18% | ALIZARINE | 0.73% | ||
| ALYCIDON | 1945 | 3.47% | DONATELLO II | 2.27% | AURORA | 3.48% | ||
| ALYDAR | 1975 | 0.54% | RAISE A NATIVE | 0.83% | SWEET TOOTH | 5.14% | ||
| APALACHEE | 1971 | 0.74% | ROUND TABLE | 0.98% | MOCCASIN | 0.79% | ||
| ASTERUS | 1923 | 1.88% | TEDDY | 1.97% | ASTRELLA | 0.85% | ||
| AUREOLE | 1950 | 1.27% | HYPERION | 3.72% | ANGELOLA | 1.62% | ||
| BACHELOR'S DOUBLE | 1906 | 4.25% | TREDENNIS | 2.64% | LADY BAWN | 2.20% | ||
| BAHRAM | 1932 | 2.08% | BLANDFORD | 3.06% | FRIAR'S DAUGHTER | 3.74% | ||
| BALDSKI | 1974 | 1.92% | NIJINSKY II | 0.54% | TOO BALD | 7.18% | ||
| BALLYMOSS | 1954 | 2.81% | MOSSBOROUGH | 4.51% | INDIAN CALL | 1.87% | ||
| BAYARDO | 1906 | 5.13% | BAY RONALD | 1.62% | GALICIA | 0.98% | ||
| BEN BRUSH | 1893 | 1.43% | BRAMBLE | 0.79% | ROSEVILLE | 0.88% | ||
| BEST TURN | 1966 | 1.13% | TURN-TO | 4.64% | SWEET CLEMENTINE | 0.73% | ||
| BIG GAME | 1939 | 3.09% | BAHRAM | 2.08% | MYROBELLA | 1.29% | ||
| BLACK TONEY | 1911 | 0.39% | PETER PAN | 0.29% | BELGRAVIA | 1.81% | ||
| BLANDFORD | 1919 | 3.06% | SWYNFORD | 1.08% | BLANCHE | 3.32% | ||
| BLENHEIM II | 1927 | 2.95% | BLANDFORD | 3.06% | MALVA | 1.75% | ||
| BLUE LARKSPUR | 1926 | 2.25% | BLACK SERVANT | 0.44% | BLOSSOM TIME | 0.73% | ||
| BLUSHING GROOM | 1974 | 1.48% | RED GOD | 1.81% | RUNAWAY BRIDE | 2.40% | ||
| BOIS ROUSSEL | 1935 | 2.10% | VATOUT | 4.13% | PLUCKY LIEGE | 1.37% | ||
| BOLD BIDDER | 1962 | 0.10% | BOLD RULER | 0.49% | HIGH BID | 1.37% | ||
| BOLD RUCKUS | 1976 | 2.87% | BOLDNESIAN | 0.93% | RAISE A RUCKUS | 1.23% | ||
| BOLD RULER | 1954 | 0.49% | NASRULLAH | 0.69% | MISS DISCO | 0.30% | ||
| BRANTOME | 1931 | 0.89% | BLANDFORD | 3.06% | VITAMINE | 1.91% | ||
| BRITISH EMPIRE | 1937 | 1.08% | COLOMBO | 3.71% | ROSE OF ENGLAND | 1.42% | ||
| BROAD BRUSH | 1983 | 4.45% | ACK ACK | 0.89% | HAY PATCHER | 0.69% | ||
| BROOMSTICK | 1901 | 0.29% | BEN BRUSH | 1.43% | ELF | 1.13% | ||
| BRULEUR | 1910 | 1.18% | CHOUBERSKI | 0.39% | BASSE TERRE | 3.03% | ||
| BUCKAROO | 1975 | 1.77% | BUCKPASSER | 1.13% | STEPPING HIGH | 0.30% | ||
| BUCKPASSER | 1963 | 1.13% | TOM FOOL | 0.63% | BUSANDA | 0.59% | ||
| BULL DOG | 1927 | 1.59% | TEDDY | 1.97% | PLUCKY LIEGE | 1.37% | ||
| BULL LEA | 1935 | 0.49% | BULL DOG | 1.59% | ROSE LEAVES | 0.93% | ||
| BUSTED | 1963 | 0.54% | CREPELLO | 0.34% | SANS LE SOU | 0.05% | ||
| CARO | 1967 | 0.45% | FORTINO II | 0.78% | CHAMBORD | 1.80% | ||
| CARSON CITY | 1987 | 1.66% | MR. PROSPECTOR | 0.44% | BLUSHING PROMISE | 0.93% | ||
| CHATEAU BOUSCAUT | 1927 | 0.39% | KIRCUBBIN | 0.73% | RAMONDIE | 2.20% | ||
| CHAUCER | 1900 | 0.78% | ST. SIMON | 0.75% | CANTERBURY PILGRIM | 1.52% | ||
| CHIEF'S CROWN | 1982 | 0.54% | DANZIG | 0.59% | SIX CROWNS | 0.98% | ||
| CICERO | 1902 | 1.56% | CYLLENE | 2.25% | GAS | 1.91% | ||
| CLARISSIMUS | 1913 | 1.93% | RADIUM | 2.16% | QUINTESSENCE | 1.18% | ||
| CODEX | 1977 | 0.65% | ARTS AND LETTERS | 0.25% | ROUNDUP ROSE | 1.13% | ||
| COLORADO | 1923 | 4.52% | PHALARIS | 1.22% | CANYON | 2.93% | ||
| CONGREVE | 1924 | 2.86% | COPYRIGHT | 4.42% | PER NOI | 3.89% | ||
| COUNT FLEET | 1940 | 0.59% | REIGH COUNT | 3.66% | QUICKLY | 0.49% | ||
| COURT MARTIAL | 1942 | 2.41% | FAIR TRIAL | 0.99% | INSTANTANEOUS | 0.15% | ||
| CREME DELA CREME | 1963 | 0.59% | OLYMPIA | 1.27% | JUDY RULLAH | 1.03% | ||
| CREPELLO | 1954 | 0.34% | DONATELLO II | 2.27% | CREPUSCULE | 0.83% | ||
| DAMASCUS | 1964 | 1.53% | SWORD DANCER | 0.93% | KERALA | 1.17% | ||
| DANZIG | 1977 | 0.59% | NORTHERN DANCER | 1.08% | PAS DE NOM | 0.30% | ||
| DARK RONALD | 1905 | 1.76% | BAY RONALD | 1.62% | DARKIE | 2.21% | ||
| DISCOVERY | 1931 | 1.27% | DISPLAY | 1.33% | ARIADNE | 0.44% | ||
| DJEBEL | 1937 | 0.25% | TOURBILLON | 0.35% | LOIKA | 4.09% | ||
| DONATELLO II | 1934 | 2.27% | BLENHEIM II | 2.95% | DELLEANA | 3.20% | ||
| DOUBLE JAY | 1944 | 1.28% | BALLADIER | 0.60% | BROOMSHOT | 0.40% | ||
| DR. FAGER | 1964 | 2.78% | ROUGH'N TUMBLE | 2.11% | ASPIDISTRA | 1.61% | ||
| EIGHT THIRTY | 1936 | 2.16% | PILATE | 2.64% | DINNER TIME | 0.44% | ||
| ELA-MANA-MOU | 1976 | 2.97% | PITCAIRN | 1.24% | ROSE BERTIN | 2.57% | ||
| EQUIPOISE | 1928 | 0.68% | PENNANT | 0.49% | SWINGING | 1.54% | ||
| EXCLUSIVE NATIVE | 1965 | 1.07% | RAISE A NATIVE | 0.83% | EXCLUSIVE | 2.84% | ||
| FAIR PLAY | 1905 | 0.64% | HASTINGS | 1.32% | FAIRY GOLD | 0.64% | ||
| FAIR TRIAL | 1932 | 0.99% | FAIRWAY | 2.80% | LADY JUROR | 0.73% | ||
| FAIRWAY | 1925 | 2.80% | PHALARIS | 1.22% | SCAPA FLOW | 3.07% | ||
| FAPPIANO | 1977 | 0.64% | MR. PROSPECTOR | 0.44% | KILLALOE | 0.49% | ||
| FORLI | 1963 | 1.91% | ARISTOPHANES | 1.61% | TREVISA | 1.23% | ||
| FOXBRIDGE | 1930 | 2.93% | FOXLAW | 3.29% | BRIDGEMOUNT | 1.97% | ||
| FULL SAIL | 1934 | 2.79% | FAIRWAY | 2.80% | FANCY FREE | 3.08% | ||
| GAINSBOROUGH | 1915 | 2.16% | BAYARDO | 5.13% | ROSEDROP | 0.59% | ||
| GALLANT MAN | 1954 | 4.16% | MIGOLI | 1.86% | MAJIDEH | 1.22% | ||
| GIANT'S CAUSEWAY | 1997 | 1.12% | STORM CAT | 0.59% | MARIAH'S STORM | 2.70% | ||
| GRAUSTARK | 1963 | 0.79% | RIBOT | 0.69% | FLOWER BOWL | 1.53% | ||
| GREY DAWN II | 1962 | 0.50% | HERBAGER | 1.47% | POLAMIA | 0.10% | ||
| GREY SOVEREIGN | 1948 | 1.99% | NASRULLAH | 0.69% | KONG | 0.64% | ||
| GUNDOMAR | 1942 | 1.71% | ALCHIMIST | 4.56% | GROSSULARIA | 5.00% | ||
| HABITAT | 1966 | 0.83% | SIR GAYLORD | 0.39% | LITTLE HUT | 0.40% | ||
| HAIL TO REASON | 1958 | 1.47% | TURN-TO | 4.64% | NOTHIRDCHANCE | 1.76% | ||
| HALO | 1969 | 1.87% | HAIL TO REASON | 1.47% | COSMAH | 0.69% | ||
| HAVRESAC II | 1915 | 12.71% | RABELAIS | 1.67% | HORS CONCOURS | 5.44% | ||
| HELIOPOLIS | 1936 | 4.91% | HYPERION | 3.72% | DRIFT | 2.35% | ||
| HERBAGER | 1956 | 1.47% | VANDALE | 0.30% | FLAGETTE | 14.71% | ||
| HIGH TOP | 1969 | 0.35% | DERRING-DO | 3.25% | CAMENAE | 0.15% | ||
| HIS MAJESTY | 1968 | 0.79% | RIBOT | 0.69% | FLOWER BOWL | 1.53% | ||
| HOIST THE FLAG | 1968 | 0.35% | TOM ROLFE | 1.43% | WAVY NAVY | 0.05% | ||
| HURRY ON | 1913 | 1.62% | MARCOVIL | 5.67% | TOUT SUITE | 2.85% | ||
| HYPERION | 1930 | 3.72% | GAINSBOROUGH | 2.16% | SELENE | 3.89% | ||
| ICECAPADE | 1969 | 0.74% | NEARCTIC | 2.85% | SHENANIGANS | 0.34% | ||
| IN REALITY | 1964 | 4.50% | INTENTIONALLY | 1.37% | MY DEAR GIRL | 0.34% | ||
| IN THE WINGS | 1986 | 0.89% | SADLER'S WELLS | 1.39% | HIGH HAWK | 0.39% | ||
| INDIAN RIDGE | 1985 | 1.08% | AHONOORA | 0.29% | HILLBROW | 0.44% | ||
| INTENTIONALLY | 1956 | 1.37% | INTENT | 0.39% | MY RECIPE | 0.05% | ||
| KEY TO THE MINT | 1969 | 0.94% | GRAUSTARK | 0.79% | KEY BRIDGE | 0.20% | ||
| KHALED | 1943 | 1.48% | HYPERION | 3.72% | ECLAIR | 1.08% | ||
| KING SALMON | 1930 | 2.17% | SALMON TROUT | 0.69% | MALVA | 1.75% | ||
| KINGMAMBO | 1990 | 1.37% | MR. PROSPECTOR | 0.44% | MIESQUE | 0.30% | ||
| KING'S BISHOP | 1969 | 0.10% | ROUND TABLE | 0.98% | SPEARFISH | 2.21% | ||
| LA FARINA | 1911 | 2.59% | SANS SOUCI II | 2.35% | MALATESTA | 1.23% | ||
| LE FABULEUX | 1961 | 0.49% | WILD RISK | 2.07% | ANGUAR | 0.49% | ||
| LOST CODE | 1984 | 0.20% | CODEX | 0.65% | LOSS OR GAIN | 1.18% | ||
| LUTHIER | 1965 | 0.88% | KLAIRON | 0.78% | FLUTE ENCHANTEE | 3.89% | ||
| LYPHARD | 1969 | 0.98% | NORTHERN DANCER | 1.08% | GOOFED | 0.15% | ||
| MAHMOUD | 1933 | 1.38% | BLENHEIM II | 2.95% | MAH MAHAL | 0.59% | ||
| MAN O' WAR | 1917 | 1.13% | FAIR PLAY | 0.64% | MAHUBAH | 1.71% | ||
| MASSINE | 1920 | 2.26% | CONSOLS | 0.35% | MAURI | 2.94% | ||
| MIDSTREAM | 1933 | 0.94% | BLANDFORD | 3.06% | MIDSUMMER | 1.08% | ||
| MIEUXCE | 1933 | 0.74% | MASSINE | 2.26% | L'OLIVETE | 0.94% | ||
| MILL REEF | 1968 | 0.30% | NEVER BEND | 0.29% | MILAN MILL | 0.50% | ||
| MOSSBOROUGH | 1947 | 4.51% | NEARCO | 1.95% | ALL MOONSHINE | 3.75% | ||
| MR. PROSPECTOR | 1970 | 0.44% | RAISE A NATIVE | 0.83% | GOLD DIGGER | 0.69% | ||
| MY BABU | 1945 | 0.20% | DJEBEL | 0.25% | PERFUME II | 5.09% | ||
| NASHUA | 1952 | 0.49% | NASRULLAH | 0.69% | SEGULA | 0.68% | ||
| NASRULLAH | 1940 | 0.69% | NEARCO | 1.95% | MUMTAZ BEGUM | 0.54% | ||
| NATIVE DANCER | 1950 | 0.05% | POLYNESIAN | 2.79% | GEISHA | 0.05% | ||
| NAVARRO | 1931 | 2.16% | MICHELANGELO | 1.14% | NUVOLONA | 2.05% | ||
| NEARCO | 1935 | 1.95% | PHAROS | 2.80% | NOGARA | 4.40% | ||
| NEVER BEND | 1960 | 0.29% | NASRULLAH | 0.69% | LALUN | 1.32% | ||
| NEVER SAY DIE | 1951 | 0.93% | NASRULLAH | 0.69% | SINGING GRASS | 0.73% | ||
| NIJINSKY II | 1967 | 0.54% | NORTHERN DANCER | 1.08% | FLAMING PAGE | 0.44% | ||
| NINISKI | 1976 | 0.39% | NIJINSKY II | 0.54% | VIRGINIA HILLS | 0.30% | ||
| NODOUBLE | 1965 | 0.34% | NOHOLME II | 1.42% | ABLA-JAY | 0.54% | ||
| NOHOLME II | 1956 | 1.42% | STAR KINGDOM | 1.47% | OCEANA | 1.63% | ||
| NORTHERN DANCER | 1961 | 1.08% | NEARCTIC | 2.85% | NATALMA | 0.60% | ||
| NUREYEV | 1977 | 2.28% | NORTHERN DANCER | 1.08% | SPECIAL | 0.34% | ||
| OLEANDER | 1924 | 2.35% | PRUNUS | 3.38% | ORCHIDEE II | 6.11% | ||
| OLYMPIA | 1946 | 1.27% | HELIOPOLIS | 4.91% | MISS DOLPHIN | 2.40% | ||
| ORBY | 1904 | 1.17% | ORME | 0.98% | RHODA B. | 0.74% | ||
| ORTELLO | 1926 | 2.76% | TEDDY | 1.97% | HOLLEBECK | 4.98% | ||
| PANORAMA | 1936 | 1.52% | SIR COSMO | 0.99% | HAPPY CLIMAX | 2.06% | ||
| PERSIAN GULF | 1940 | 0.93% | BAHRAM | 2.08% | DOUBLE LIFE | 1.66% | ||
| PETER PAN | 1904 | 0.29% | COMMANDO | 2.43% | CINDERELLA | 1.43% | ||
| PETITION | 1944 | 1.23% | FAIR TRIAL | 0.99% | ART PAPER | 2.08% | ||
| PHALARIS | 1913 | 1.22% | POLYMELUS | 1.42% | BROMUS | 8.16% | ||
| PHARIS II | 1936 | 2.96% | PHAROS | 2.80% | CARISSIMA | 2.16% | ||
| PHAROS | 1920 | 2.80% | PHALARIS | 1.22% | SCAPA FLOW | 3.07% | ||
| PIA STAR | 1961 | 1.65% | OLYMPIA | 1.27% | INQUISITIVE | 0.20% | ||
| PILATE | 1928 | 2.64% | FRIAR ROCK | 3.64% | HERODIAS | 1.22% | ||
| PLEASANT COLONY | 1978 | 0.49% | HIS MAJESTY | 0.79% | SUN COLONY | 0.39% | ||
| POLYNESIAN | 1942 | 2.79% | UNBREAKABLE | 3.40% | BLACK POLLY | 3.32% | ||
| POMPEY | 1923 | 1.96% | SUN BRIAR | 1.27% | CLEOPATRA | 2.47% | ||
| PRECIPITATION | 1933 | 0.88% | HURRY ON | 1.62% | DOUBLE LIFE | 1.66% | ||
| PRETENSE | 1963 | 3.45% | ENDEAVOUR II | 1.91% | IMITATION | 4.64% | ||
| PRINCE BIO | 1941 | 0.94% | PRINCE ROSE | 1.50% | BIOLOGIE | 1.86% | ||
| PRINCE CHEVALIER | 1943 | 0.94% | PRINCE ROSE | 1.50% | CHEVALERIE | 1.08% | ||
| PRINCE JOHN | 1953 | 0.20% | PRINCEQUILLO | 1.10% | NOT AFRAID | 1.03% | ||
| PRINCE ROSE | 1940 | 1.50% | ROSE PRINCE | 1.14% | INDOLENCE | 0.77% | ||
| PRINCEQUILLO | 1928 | 1.10% | PRINCE ROSE | 1.50% | COSQUILLA | 1.50% | ||
| PROMISED LAND | 1954 | 0.20% | PALESTINIAN | 1.48% | MAHMOUDESS | 1.14% | ||
| RABELAIS | 1900 | 1.67% | ST. SIMON | 0.75% | SATIRICAL | 3.23% | ||
| RAINBOW QUEST | 1981 | 1.18% | BLUSHING GROOM | 1.48% | I WILL FOLLOW | 0.93% | ||
| RAISE A NATIVE | 1961 | 0.83% | NATIVE DANCER | 0.05% | RAISE YOU | 0.54% | ||
| RELIANCE II | 1962 | 0.05% | TANTIEME | 0.44% | RELANCE III | 0.40% | ||
| RELKO | 1960 | 0.10% | TANERKO | 2.11% | RELANCE III | 0.40% | ||
| REVIEWER | 1966 | 1.71% | BOLD RULER | 0.49% | BROADWAY | 1.76% | ||
| RIBOT | 1952 | 0.69% | TENERANI | 1.87% | ROMANELLA | 2.46% | ||
| RIGHT ROYAL | 1958 | 1.37% | OWEN TUDOR | 3.72% | BASTIA | 3.28% | ||
| RIVERMAN | 1969 | 0.69% | NEVER BEND | 0.29% | RIVER LADY | 0.29% | ||
| ROBERTO | 1969 | 3.15% | HAIL TO REASON | 1.47% | BRAMALEA | 1.27% | ||
| ROCK SAND | 1900 | 4.21% | SAINFOIN | 6.57% | ROQUEBRUNE | 2.16% | ||
| ROMAN | 1937 | 0.20% | SIR GALLAHAD III | 1.59% | BUCKUP | 0.15% | ||
| ROUGH'N TUMBLE | 1948 | 2.11% | FREE FOR ALL | 0.89% | ROUSED | 0.34% | ||
| ROUND TABLE | 1954 | 0.98% | PRINCEQUILLO | 1.10% | KNIGHT'S DAUGHTER | 0.94% | ||
| ROYAL ACADEMY | 1987 | 1.12% | NIJINSKY II | 0.54% | CRIMSON SAINT | 0.44% | ||
| ROYAL CHARGER | 1942 | 0.74% | NEARCO | 1.95% | SUN PRINCESS | 2.00% | ||
| RUN THE GANTLET | 1968 | 2.31% | TOM ROLFE | 1.43% | FIRST FEATHER | 0.10% | ||
| SADLER'S WELLS | 1981 | 1.39% | NORTHERN DANCER | 1.08% | FAIRY BRIDGE | 0.39% | ||
| SARDANAPALE | 1911 | 0.24% | PRESTIGE | 1.12% | GEMMA | 1.13% | ||
| SEA-BIRD | 1962 | 0.49% | DAN CUPID | 4.45% | SICALADE | 1.95% | ||
| SEATTLE SLEW | 1974 | 1.57% | BOLD REASONING | 1.53% | MY CHARMER | 2.15% | ||
| SECRETARIAT | 1970 | 0.24% | BOLD RULER | 0.49% | SOMETHINGROYAL | 0.34% | ||
| SHARPEN UP | 1969 | 1.49% | ATAN | 0.88% | ROCCHETTA | 0.40% | ||
| SHIRLEY HEIGHTS | 1975 | 0.30% | MILL REEF | 0.30% | HARDIEMMA | 0.20% | ||
| SICAMBRE | 1948 | 1.07% | PRINCE BIO | 0.94% | SIF | 9.10% | ||
| SIDERAL | 1948 | 1.13% | SEDUCTOR | 3.09% | STARLING II | 1.18% | ||
| SIR COSMO | 1926 | 0.99% | THE BOSS | 1.03% | AYN HALI | 0.54% | ||
| SIR GALLAHAD III | 1920 | 1.59% | TEDDY | 1.97% | PLUCKY LIEGE | 1.37% | ||
| SIR GAYLORD | 1959 | 0.39% | TURN-TO | 4.64% | SOMETHINGROYAL | 0.34% | ||
| SIR IVOR | 1965 | 1.22% | SIR GAYLORD | 0.39% | ATTICA | 0.59% | ||
| SMART STRIKE | 1992 | 0.79% | MR. PROSPECTOR | 0.44% | CLASSY 'N SMART | 0.69% | ||
| SOLARIO | 1922 | 2.47% | GAINSBOROUGH | 2.16% | SUN WORSHIP | 2.94% | ||
| SON-IN-LAW | 1911 | 2.41% | DARK RONALD | 1.76% | MOTHER-IN-LAW | 1.66% | ||
| SPEAK JOHN | 1958 | 0.00% | PRINCE JOHN | 0.20% | NUIT DE FOLIES | 0.29% | ||
| SPEARMINT | 1903 | 2.84% | CARBINE | 4.57% | MAID OF THE MINT | 6.02% | ||
| SPY SONG | 1943 | 3.17% | BALLADIER | 0.60% | MATA HARI | 2.25% | ||
| STAGE DOOR JOHNNY | 1965 | 0.10% | PRINCE JOHN | 0.20% | PEROXIDE BLONDE | 0.34% | ||
| STAR KINGDOM | 1946 | 1.47% | STARDUST | 2.36% | IMPROMPTU | 1.08% | ||
| STAR SHOOT | 1898 | 1.71% | ISINGLASS | 2.95% | ASTROLOGY | 1.14% | ||
| SUNNY BOY | 1944 | 4.05% | JOCK | 2.26% | FILLE DE SOLEIL | 1.03% | ||
| SUNSTAR | 1908 | 2.40% | SUNDRIDGE | 3.33% | DORIS | 1.49% | ||
| SWEEP | 1907 | 1.96% | BEN BRUSH | 1.43% | PINK DOMINO | 0.84% | ||
| SWYNFORD | 1907 | 1.08% | JOHN O'GAUNT | 1.23% | CANTERBURY PILGRIM | 1.52% | ||
| T.V. LARK | 1957 | 1.27% | INDIAN HEMP | 2.20% | MISS LARKSFLY | 0.35% | ||
| TANTIEME | 1947 | 0.44% | DEUX POUR CENT | 0.68% | TERKA | 0.94% | ||
| TEDDY | 1913 | 1.97% | AJAX | 1.38% | RONDEAU | 1.71% | ||
| THE TETRARCH | 1911 | 4.35% | ROI HERODE | 2.61% | VAHREN | 4.60% | ||
| TICINO | 1939 | 0.93% | ATHANASIUS | 0.74% | TERRA | 1.82% | ||
| TOM FOOL | 1949 | 0.63% | MENOW | 1.92% | GAGA | 0.34% | ||
| TOM ROLFE | 1962 | 1.43% | RIBOT | 0.69% | POCAHONTAS | 0.93% | ||
| TOURBILLON | 1928 | 0.35% | KSAR | 7.13% | DURBAN | 2.58% | ||
| TRACERY | 1909 | 3.05% | ROCK SAND | 4.21% | TOPIARY | 1.51% | ||
| TRAGHETTO | 1942 | 1.87% | CAVALIERE D'ARPINO | 2.75% | TALMA | 1.28% | ||
| TUDOR MINSTREL | 1944 | 2.06% | OWEN TUDOR | 3.72% | SANSONNET | 0.35% | ||
| TURN-TO | 1951 | 4.64% | ROYAL CHARGER | 0.74% | SOURCE SUCREE | 1.67% | ||
| ULTIMUS | 1906 | 14.32% | COMMANDO | 2.43% | RUNNING STREAM | 1.32% | ||
| UNBRIDLED | 1987 | 1.23% | FAPPIANO | 0.64% | GANA FACIL | 0.00% | ||
| VAGUELY NOBLE | 1965 | 3.95% | VIENNA | 1.23% | NOBLE LASSIE | 1.27% | ||
| VANDALE | 1943 | 0.30% | PLASSY | 2.41% | VANILLE | 1.97% | ||
| VATELLOR | 1933 | 0.74% | VATOUT | 4.13% | LADY ELINOR | 1.47% | ||
| VATOUT | 1926 | 4.13% | PRINCE CHIMAY | 1.86% | VASHTI | 1.71% | ||
| VIEUX MANOIR | 1947 | 0.73% | BRANTOME | 0.89% | VIEILLE MAISON | 0.29% | ||
| WAR ADMIRAL | 1934 | 0.29% | MAN O' WAR | 1.13% | BRUSHUP | 0.10% | ||
| WHAT A PLEASURE | 1965 | 2.90% | BOLD RULER | 0.49% | GREY FLIGHT | 0.44% | ||
| WILD RISK | 1940 | 2.07% | RIALTO | 1.19% | WILD VIOLET | 1.37% | ||
| WORDEN | 1949 | 0.35% | WILD RISK | 2.07% | SANS TARES | 1.03% |